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Doctor of Philosophy (Ph.D) in Evolution, Ecology and Organismal Biology



What will I learn?

Specializations for graduate studies are Evolutionary Biology, Ecology and Physiology. The Department's research in Evolutionary Biology encompasses evolutionary ecology and population genetics, behavioral ecology, molecular systematics, physiological ecology, and evolutionary physiology. We are also concerned with the application of knowledge from these disciplines to problems in biological control and conservation biology. Our Physiologists work in the following areas: physiological ecology (physiological adaptations of animals to their environments), comparative and evolutionary physiology (evolutionary changes in physiological traits), and regulatory physiology (especially mechanisms of water balance and thermoregulation).

The Ph.D. degree requires demonstration of broad knowledge of Ecology, Evolutionary Biology, and/or Organismal Biology, and substantive ability and accomplishment in original research.

Which department am I in?

College of Natural and Agricultural Sciences

Study options

Full Time (15 quarters)

Tuition fees
US$32,287.86 per year
Application deadline

Expected March, June 2023

Start date

Expected March, June, September 2023


College of Natural and Agricultural Sciences

Geology Building, Room 2258,

900 University Avenue,



92521, United States

Entry requirements

For students from United States

To be admitted to graduate status, the basic requirement is a bachelor's degree in Physics or its equivalent from an accredited institution. This degree must represent the completion of a program equivalent both in the distribution of academic subject matter and scholarship achievement comparable to that offered at the University of California. Qualified applicants are expected to have a junior/senior GPA of 3.25 or higher.

For international students

A B.A. or B.S. degree in Biology or a related discipline. Majors in other scientific fields are acceptable if basic preparation in biology, chemistry (organic and inorganic), physics, and mathematics are deemed adequate. This should include calculus, inorganic and organic chemistry, and physics.

English Language Requirement: TOEFL - The minimum acceptable scores are: 550 for the revised TOEFL paper-delivered test and 80 for the TOEFL iBT; IELTS - The minimum acceptable overall score is 7 with no score less than 6 on any individual component.

Application deadline: Summer - March 1st; Fall - June 1st; Winter - September 1st; Spring - December 1st.

*There may be different IELTS requirements depending on your chosen course.


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